Manny (voice)

Ray Romano

Sid

John Leguizamo

Sid

John Leguizamo

Diego

Denis Leary

Ellie (voice)

Queen Latifah

Crash

Seann William Scott

Scrat

Chris Wedge

Eddie (voice)

Josh Peck

Fast Tony

Jay Leno

Lone Gunslinger Vulture

Will Arnett

Grandpa Molehog

Jason Fricchione

Dung Beetle Dad

Peter Ackerman

Freaky Female

Denny Dillon

Mr. Start

Joseph Bologna

Elk Boy

Jack Crocicchia

Glypto Boy Billy / Beaver Girl (voice)

Caitlin Rose Anderson

Glypto Boy Billy/Beaver Girl

Caitlin Rose Anderson

Rhino Boy / Beaver Boy (voice)

Connor Anderson

Rhino Boy/Beaver Boy

Connor Anderson

Condor Chick

Peter DeSève

Elk Boy (voice)

Jack Crocicchia

Start Girl

Nicole DeFelice

Condor Chick (voice)

Peter DeSève

Additional Voices (voice)

Ariel Winter

Diatryma Mom

Debi Derryberry

Start Dad

Marshall Efron

Female Mini Sloth (voice)

Clea Lewis

Aardvark Dad (voice)

Stephen Root

Stu/Male Ox

Tom Fahn

Start Girl (voice)

Nicole DeFelice

Traffic Vulture

James Edmund Godwin

Beaver Dad

George Jacobs

Shovelmouth Boy

Jansen Panettiere

Start Boy No. 1

Gregory Romano

Traffic Vulture (voice)

James Edmund Godwin

Start Boy No. 2

Matthew Romano

Glyptodon (voice)

Brian Scott McFadden

Dodo

Carlos Saldanha

Molehog Kid No. 1

Manoela Scarpa Saldanha

Start Boy 1 (voice)

Gregory Romano

Molehog Kid No. 2

Sofia Scarpa Saldanha

Start Boy 2 (voice)

Matt Romano

Diego's Bird Girl

Reyna Shaskan

Freaky Male

James Sie

Aardvark Mom

Cindy Slattery

Female Ox

Mindy Sterling

Diego's Bird Girl (voice)

Reyna Shaskan

James

Alex Sullivan

Mrs. Start

Renée Taylor

Cholly

Alan Tudyk

Arne Elsholtz

James (voice)

Alex Sullivan

Otto Waalkes

Thomas Fritsch

Rainer Fritzsche

Additional Voices (voice)

Claudia Besso

Daniela Hoffmann

Additional Voices (voice)

Madeleine Martin

Julien Haggége

Additional Voices (voice)

Sherry Lynn

Additional Voices (voice)

Mickie McGowan

Lutz Mackensy

Additional Voices (voice)

Emily Anderson

Oliver Stritzel

Marlon Flechtner

Additional Voices (voice)

Jack Angel

Lennart Flechtner

Additional Voices (voice)

Shane Baumel

Daniela Reidies

Additional Voices (voice)

Bob Bergen

Additional Voices (voice)

Kyle J. Bogert

Additional Voices (voice)

Rodger Bumpass

Brian McFadden

Matt Romano

Additional Voices (voice)

Paul Butcher

Otto Jespersen

Additional Voices (voice)

Blanca Camacho

Additional Voices (voice)

John Cygan

Dagfinn Lyngbø

Additional Voices (voice)

Jennifer Darling

Sven Nordin

Additional Voices (voice)

Olivia DeLaurentis

Linn Skåber

Espen Sandvik

Additional Voices (voice)

Harrison Fahn

Additional Voices (voice)

Bill Farmer

Jan Gunnar Røise

Jan Erik Larssen

Additional Voices (voice)

Crispin Freeman

Robert Gustafsson

Additional Voices (voice)

Tim Kirkpatrick

Additional Voices (voice)

David H. Kramer

Björn Granath

Reine Brynolfsson

Additional Voices (voice)

Danny Mann

Additional Voices (voice)

Laura Marano

Eva Röse

Göran Gillinger

Additional Voices (voice)

Matt McCarthy

Andreas Nilsson

Additional Voices (voice)

Laraine Newman

Additional Voices (voice)

Jan Rabson

Johan Ulveson

Anna Nordell

Additional Voices (voice)

Zack Shada

Lars Amble

Additional Voices (voice)

Thomas Sharkey

Additional Voices (voice)

Ross Simanteris

Jens Hultén

Lars Dejert

Additional Voices (voice)

Rachel Stand

Emil Smedius

Additional Voices (voice)

Spencer Lacey Ganus

Simon Sjöquist

Additional Voices (voice)

Jess Harnell

Jonathan Selander

Additional Voices (voice)

Gérard Lanvin

Eddie Hultén

Additional Voices (voice)

Élie Semoun

Mikaela Ardai Jennefors

Additional Voices (voice)

Vincent Cassel

Norea Sjöquist

Additional Voices (voice)

Christophe Dechavanne

Göran Engman

Additional Voices (voice)

Manu Payet

Additional Voices (voice)

Florian Gazan

Additional Voices (voice)

Bruno Guillon

Armelle Gallaud

Christophe Dechavanne

Stefan Gossler

David Hellwig

Actor

Director

Carlos Saldanha